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NOX4, acting as a source of cellular superoxide anions, not only enhances the expression of CPT1A but also increases the production of mtROS and self-derived ROS, ultimately activating the NLRP3 inflammasome [127, 128]

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Cell culture and stimulation Normal human epidermal keratinocytes (NHEKs) were purchased from Takara Bio (Shiga, Japan) and cultured in Keratinocyte Basal Medium (KBM) overnight prior to human recombinant rIL-22 stimulation (200 ng ml 1 , R&D Systems, Minneapolis, MN)