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caldus strains grown for 7 days without heavy metals were supplemented with CuSO 4 (final concentration: 5 mM) or ZnSO 4 (final concentration: 40 mM), respectively, incubated at 40C and shaken at 150 rpm

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10.1016/j.clinbiochem.2012.01.023 34

The first and rate-limiting step is the synthesis of a dipeptide, catalyzed by -glutamylcysteine synthase

Therefore, the exogenous (dietary) antioxidants vitamin E and carotenoids are essential to protect cell membranes and the membranes surrounding organelles (e.g., mitochondria) from radical-mediated damage