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Acknowledgments This work was supported by the Institut National de la Recherche en Agronomie, the Centre National de la Recherche Scientifique, the Cte dAzur University, and the French Government (National Research Agency, ANR) through the LABEX SIGNALIFE program (reference # ABR-11-LABX-0028-01) and the STAYPINK project (ANR-15-CE20-0005)