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Cells were harvested by centrifugation at 3000 rcf for 15 min, washed with ice cold PBS and resuspended in lysis buffer [25 mm HEPES-KOH (pH 7.4), 400 mm KCl, 10% glycerol, 1% Triton X-100, 25 mm imidazole, and 1 mm dithiothreitol (DTT)]

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