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Acknowledgements We thank the members of the Nudler and Littman laboratories for their discussions and review of the manuscript

Cytosolic and mitochondrial pools of malate and OAA are connected by two transporters the malate-2OG transporter (Slc25a11) and the glutamate-aspartate transporter (Slc1a3) and by the interconversion of aspartate and glutamate into OAA and 2OG by the cytosolic and mitochondrial isoforms of the glutamate-oxaloacetate transaminase, Got1 and Got2, respectively