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Zhang et al, 2014) reported that limonene participates in the mitochondrial pathway of apoptosis, increases the expression of BAX protein, and decreases BCL2 protein expression, thus confirming its pro-apoptotic activity

Furthermore, increased strand breaks generation caused activation of poly(ADP-ribose) polymerase (PARP), a zinc finger DNA-binding protein, known to deplete intracellular NAD + reserves as well as energy stores, thus ultimately resulting in cell death (429)

Similarly, under UVA irradiation, glutathione amino acid precursor treatment resulted in lower levels of oxidative DNA lesions, as indicated by 8-OHdG staining