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1 Introduction Since the pre-industrial period, atmospheric CO 2 concentrations have increased from 277 to 414 ppm in 2021, and they have continued to increase by approximately 2.4 ppm per year over the last decade, reaching 1.070 ppm until the end of the century (Friedlingstein et al., 2 and heat (Caldeira and Wickett, Ocean acidification and warming are critical threats to coastal ecosystems (Boyd et al., 2023