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Under oxygen enrichment, the primary function of glycolysis is to produce intermediate metabolites that provide source materials for biosynthetic pathways, whereas under hypoxia, the primary role of glycolysis is to provide ATP for cell survival (37)

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It is deliberately membrane-permeable, which is what lets it get inside cells and act on an intracellular enzyme

Weber, R

Nature 613 , 355364 (2023)

Berberine metabolically modulates lipid profiles by reducing intracellular cholesterol synthesis and promoting cholesterol efflux