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24 The early work of George and Mildred Burr established that the C18 polyunsaturated long-chain fatty acids (PUFAs) linoleic acid (LA, 18:2 6) and -linolenic acid (ALA, 18:3 3) are dietary essential fatty acids (EFAs) 5 because humans and most mammals lack the desaturase enzymes that can introduce an 6 (n-6) or 3 (n-3) double bond beyond carbons 9 and 10

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