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In contrast, the lifespan of mitochondrial electron transport chain deficient isp-1 mutant and raga-1 mutants, in which mTOR complex 1 activity is inhibited, remains unaffected by changes in the media (Figure 4C and D), indicating that the lifespan extension effects induced by mitochondria deficiency and TOR inhibition are resilient to environmental factors that interfere with metformin action in aging

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