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Acknowledgements We thank the University of Pennsylvania Diabetes Research Center (DRC) for use of the RIA Biomarker Core (P30-DK19525), and the University of Pennsylvania Rodent Metabolic Phenotyping Core (RRID:SCR_022427), supported in part by NIH grant S10-OD025098, for use of the EchoMRI to perform body composition measurements and for performing metabolic exercise performance experiments