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Comprehensive genomic analyses have revealed a distinct tobacco-related mutational signature in liver tumors of smokers, characterized by cytosine-to-adenine base substitutions, dinucleotide alterations, and guanine adduct formation, patterns resembling those observed in lung and head-and-neck cancers [3840]

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Human bronchial epithelial cells (HBECs, ZQ0001, ScienCell, San Diego, California, USA) were cultured in Keratinocyte Medium (#2111, ScienCell, San Diego, California, USA), supplemented with 100 U/ml penicillin, 100 g/ml streptomycin and 1 keratinocyte growth supplement (#2162, ScienCell, San Diego, California, USA)

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