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The released Fe 3+ is then reduced to Fe 2+ by six-transmembrane epithelial antigen of prostate 3 (STEAP3) and exported from the endosome into the cytoplasm by divalent metal transporter1 (DMT1)

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Methods to measure brain aging have used MRI-based brain volume and functional connectivity measurements, which are costly and do not provide molecular insights 16 , or have required tissue samples, which prevents their application in living persons 17

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